Article,

Population Structure in Toads

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The American Naturalist, 77 (773): 563-568 (1943)

Abstract

(1) Data on population structure in toads, chiefly Bufo americanus, were gathered in northeastern Oklahoma in the spring of 1941. (2) In the area studied, B. americanus males greatly outnumbered females in the breeding ponds (6:1 considering all breeding aggregations where both male and female toads were present). (3) Mixed aggregations of B. americanus and B. woodhousii occur. (4) The average size of 15 breeding aggregations of B. americanus was found to be 19.3 toads (16.6 males and 2.6 females). (5) The male chorus daily loses and gains members. Male toads which do not find mates may continue calling in the same pond or may migrate to another pond and call there.

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