Angluin et al. proved that population protocols compute exactly the predicates definable in Presburger arithmetic (PA), the first-order theory of addition. As part of this result, they presented a procedure that translates any formula $\varphi$ of quantifier-free PA with remainder predicates (which has the same expressive power as full PA) into a population protocol with $2^O(poly(|\varphi|))$ states that computes $\varphi$. More precisely, the number of states of the protocol is exponential in both the bit length of the largest coefficient in the formula, and the number of nodes of its syntax tree. In this paper, we prove that every formula $\varphi$ of quantifier-free PA with remainder predicates is computable by a leaderless population protocol with $O(poly(|\varphi|))$ states. Our result shows that, contrary to the case of time complexity, where protocols with leaders can be exponentially faster than leaderless protocols, protocols with and without leaders have both polynomial state complexity. Our proof is based on several new constructions, which may be of independent interest. Given a formula $\varphi$ of quantifier-free PA with remainder predicates, a first construction produces a succinct protocol (with $O(|\varphi|^3)$ leaders) that computes $\varphi$; this completes the work initiated in STACS'18, where we constructed such protocols for a fragment of PA. For large enough inputs, we can get rid of these leaders. If the input is not large enough, then it is small, and we design another construction producing a succinct protocol with one leader that computes $\varphi$. Our last construction gets rid of this leader for small inputs.

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