Abstract
The genetic variability in a subdivided population of finite size was in-
vestigated in terms of the effective number of alleles maintained. The population
structures used in the investigation are the island model and the stepping stone
model. For these models, exact formulas or approximation formulas which give
the probability that two randomly chosen homologous genes, one from each of
two specified colonies (or both from the same colony) are identical by descent
were obtained. It is shown that the genetic variability in the whole population
with stepping stone structure is known if the mutation rate and the variability in
a single colony is known, without knowing the migration rate. The validity of
the analyses was checked by simulation.
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