Abstract
We study a simple model of eco-systems described by the replicator
dynamics
equationeq:def
n_i(t)n_i(t)=f(n_i(t))+1N\sum_\mu=1^
N Q^\mu(t)\xi_i^\mu+\sum_j J_ij n_j(t)-\nu(t),
equation
where $n_i(t)$ dentes the abandance of species $i=1,,N$ at time
$t$, $Q^\mu(t)$ the abundance of resource $\mu=1,,P=N$
and $\xi_i^\mu$ consumption of resource $\mu$ by species $i$.
We assume the diversity $P$ of the resources is comparable to the
number of species $N$ , i.e. $\alpha=O(1)$. Although it is
similar to the model proposed in (De Martino and Marsili, 2006), but
we newly consider effects by random inter-species interactions $\J_
ij\$ obeying a gaussian distribution with mean zero, variance $w/N
$ and correlation $J_ijJ_ji\rangle_J=\Gamma w/N$. By
the parameter $\Gamma$, the model represents a variety of ecological
interactions, for example,
antisymmetric interaction corresponds to prey-predator interactions
in a food web.
The first term $f(n_i(t))$ represents co-operation pressure or
carrying capacities in a Lotka-Volterra system. The last term $\nu(t)
$ is a Lagrange multiplier to keep normalisation. Time evolution of
the resource $\mu$ is given by
equation
Q^\mu(t)=Q_0^\mu-\sum_j=1^N \xi_j^n_j(t)
equation
where $Q_0^\mu$ is steady resource abundance in absence of species,
which obeys a Gaussian
distribution with mean $P$ and variance $\sigma^2 P$. To study this
system we apply the generating
functional (GF) method based on path-integrals,which gives us a
powerful tool to analyze systems
with asymmetric disordered interactions. For such a system, there is
no Lyapnov function and static
approach like replica method are inapplicable. The GF allows one to
formulate a closed theory for
macroscopic order parameters of the system, such as the correlation
and response functions, which are
determined as self-consistent averages over an effective single-
species stochastic process, subject to
non-Markovian interaction and coloured noise. From the effective
process, we can discuss the stability of
this eco-system and the composition of surviving species as a
function of the model parameters. We
focused on the effects of variability of resources, direct
interaction between species, co-operation
pressure and dilution on the stability and the diversity of the
ecosystem.
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